The 2nd Gds Bde MG Coy was formed between 01 September and 19 September, 1915.
| Brigade attached to: | 2nd Guards Brigade |
| Division attached to: | Guards Division |
| Formed from the: | Machine Gun Section of 3rd Bn. The Grenadier Guards |
| Machine Gun Section of 1st Bn. The Coldstream Guards | |
| Machine Gun Section of 1st Bn. The Scots Guards | |
| Machine Gun Section of 2nd Bn. The Irish Guards |
The 2nd Gds Bde MG Coy was amalgamated with the 1st, 3rd, and 4th to form the 4th Bn, MG Guards on the 01 March 1918. On 10 May 1918, by authority of Royal Warrant, the Guards Machine Gun Regiment was formed and the designation of the 4th Bn became the 4th Bn, Guards MG Regt.
As a unit of the Guards Division, it will have taken part in the following battles and engagements.
| 1915 | |
| 26 September to 08 October | Battle of Loos [XI. Corps, First Army]. |
| 16 to 19 October | Hohenzollern Redoubt [XI. Corps, First Army]. |
| 1916 | |
| BATTLES OF THE SOMME | |
| 15 to 16 September | Battle of Flers-Courcellete [XIV. Corps, Fourth Army]. |
| 20 to 22 September | |
| 25 to 28 September | Battle of Morval [XIV. Corps, Fourth Army]. |
| 25 September | Capture of Lesboeufs. |
| 1917 | |
| 14 to 24 March | German Retreat to the Hindenburg Line [XIV. Corps, Fourth Army]. |
| BATTLES OF YPRES | |
| 31 July to 02 August | Battle of Pilckem Ridge [XIV. Corps, Fifth Army]. |
| 09 October | Battle of Poelcappelle [XIV. Corps, Fifth Army]. |
| 12 October | First Battle of Passchendaele [XIV. Corps, Fifth Army]. |
| BATTLE OF CAMBRAI | |
| 24 to 28 November | Capture of Bourlon Wood [IV. Corps, Third Army]. |
| 30 November to 03 December | German Counter-Attacks [III. Corps, Third Army]. |
| 30 November | Recapture of Gouzeaucourt. |
