A Machine Gun Company was attached to each Infantry Brigade and their subsequent Division.
| Brigade attached to: | 1st Brigade |
| Division attached to: | 1st Division |
| Formed from the: | Machine Gun Section of 1st Bn. The Black Watch |
| Machine Gun Section of 1st Bn. The Cameron Highlanders | |
| Machine Gun Section of 10th Bn. The Gloucestershire Regiment | |
| Machine Gun Section of 8th Bn. The Royal Berkshire Regiment |
The 1st Bde. M.G. Coy. was formed on the 26 January 1916 from the Machine Gun Sections of the listed infantry battalions.
On 28 February 1918, it was amalgamated with the 2nd, 3rd and 216th Machine Gun Companies to form No. 1 Battalion, Machine Gun Corps.
As a unit of the 1st Infantry Division, it will have taken part in the following battles and engagements.
| 1916 | |
| BATTLES OF THE SOMME | |
| 14 to 17 July | Battle of Bazentin Ridge [III Corps, Fourth Army] |
| 23 to 26 July | Battle of Pozieres Ridge [III Corps, Fourth Army] |
| 15 August to 3 September | |
| 15 to 22 September | Battle of Flers-Courcelette [III Corps, Fourth Army] |
| 25 to 28 September | Battle of Morval [III Corps, Fourth Army] |
| 1917 | |
| 14 to 21 March | German Retreat to the Hindenburg Line [III Corps, Fourth Army] |
| 21 June to 20 October | Operations on the Flanders Coast |
| 10 July | Defence of Nieuport [XV Corps, Fourth Army] |
| BATTLE OF YPRES | |
| 5 to 10 November | Second Battle of Passchendaele [II Corps, Second Army] |
| 1918 | |
| BATTLES OF THE LYS | |
| 9 to 11 April | Battle of Estaires [I Corps, First Army] |
| 15 April | Battle of Hazebrouck (3rd Bde.) [under 55th Division, XI Corps, First Army] |
| 18 and 19 April | Battle of Bethune [I Corps, First Army] |
| THE ADVANCE TO VICTORY | |
| SECOND BATTLES OF ARRAS | |
| 2 and 3 September | Battle of the Drocourt-Queant Line [Cdn. Corps, First Army] |
| BATTLES OF THE HINDERBURG LINE | |
| 18 September | Battle of Epehy [IX Corps, Fourth Army] |
| 29 September to 2 October | Battle of the St. Quentin Canal [IX Corps, Fourth Army] |
| 3 to 5 October | Battle of the Beaurevoir Line [IX Corps, Fourth Army] |
| THE FINAL ADVANCE IN PICARDY | |
| 17 to 25 October | Battle of the Selle [IX Corps, Fourth Army] |
| 4 November | Battle of the Sambre [IX Corps, Fourth Army] |
| 4 November | Passage of the Sambre-Oise Canal |
