A Machine Gun Company was attached to each Infantry Brigade and their subsequent Division.
| Brigade attached to: | 5th Brigade |
| Division attached to: | 2nd Division |
| Formed from the Machine Gun Sections of: | 2nd Bn. Oxfordshire & Buckinghamshire Light Infantry 2nd Bn. Highland Light Infantry 17th Bn. Royal Fusiliers 24th Bn. Royal Fusiliers |
The 5th Bde. M.G. Coy. was formed on the 01 January 1916 from the Machine Gun Sections of the listed infantry battalions.
As a unit of the 2nd Division, it will have taken part in the following battles and engagements.
| 1916 | |
| 25 July to 09 August | Battle of Delville Wood [XIII. Corps, Fourth Army]. |
| 17 and 28 July | Capture and consilidation of Delville Wood. |
| 08 and 09 August | Attack of Waterlot Farm – Guillemont. |
| 13 to 16 November | Battle of the Ancre [V. Corps, Fifth Army]. |
| 1917 | |
| 11 January to 13 March | Operations on the Ancre [II. Corps, Fifth Army]. |
| 17 and 18 February | Actions of Miraumont. |
| 25 February to 02 March | Capture of the Thilloys. |
| 10 March | Capture of Grevillers Trench (near Iries). |
| 14 to 19 March | German Retreat to the Hindenburg Line [II. Corps, Fifth Army]. |
| BATTLES OF ARRAS | |
| 12 to 14 April | Battle of Vimy Ridge [XIII. Corps, First Army]. |
| 28 and 29 April | Battle of Arleux [XIII. Corps, Fifth Army]. |
| 03 May | Third Battle of the Scarpe [XIII. Corps, Fifth Army]. |
| BATTLE OF CAMBRAI | |
| 27 and 28 November | Capture of Bourlon Wood [IV. Corps, Third Army]. |
| 30 November to 03 December | German Counter Attacks [IV. Corps, handed over on 1/12 to V. Corps, Third Army]. |
On 04 March 1918, it was amalgamated with the 6th, 99th and 242nd Machine Gun Companies to form No. 2 Battalion, Machine Gun Corps.
